Yay after reading 7 pages i found what i was looking for!
Nature is chaotic thats true, but i would go even further. (I know this probably would be for a much later update, but i can dream right?
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Adding bigger treats that are almost unavoidable for your ants, treats you can't conquer (looking at freeplay for example, you can overrule the entire map if you play correctly, no challenges to stay alive when you conquered the early games.) BUT adding treats, that are handled without proper care, could destroy the colony. For example, when it starts to rain, ants need to get back into their colony in time... otherwise they will all die (for example, i'm exagerating to make my point, don't take it tooo serious).
Lightning hitting plants or trees and leave them burning, if ants pass through, they also die. Earthquakes that collapses the underground... making rooms that are empty... filled randomly again. If dirt falls on eggs or storageplace, they get destroyed...
Ant-eaters like fireflame said. Just passes through randomly... starts licking with his tongue at the entrance to find something to eat.. and he can eats lots if they keep coming.
and now i'm going to say something i know is for way way way more in the future and i don't think this will ever make it in this game but... for example a giant foot walking or passing by on the map leaving destruction behind (Yes, from a human) A shadow appears, gets bigger and bam, everything underneed, dies.. The positive side of this.. You can combine it also with a cookie that falls randomly through the sky.. attracting other species also ofcourse.
Same as a human trying to destroy a nest, pouring toxic over the map or at some places... ants that are infected can still survive for like 3 minutes but then die afterwards.
i'm kinda brainstorming here, i really think some ideas would make the game better, i know it can take some time when we hit this tho. Nevertheless would be a cool extra option at free play or in some game modes.